♂ (dsf), Mae Hong Son, Thailand.		♀ (dsf), Chiang Mai, Thailand.
♂ (wsf), Chiang Mai, Thailand.		♀ (wsf), Tak, Thailand.

ON : Isoteinon atkinsoni Moore, 1878 OD : Proc. zool. Soc. Lond. 1878(3):693, pl.45, fig.10,♂(Un). TL : Darjiling. (♂) Distribution : Sikkim to Myanmar, Thailand, Laos, Vietnam.

Isoteinon khasianus Moore, 1878 : Proc. zool. Soc. Lond. 1878(3):693. TL. Khasia Hills. (♂, NHML)
Isoteinon subtestaceus Moore, 1879 : Proc. zool. Soc. Lond. 1878(4):844. TL. Ahwown, Tenasserim. (♂, NHML)
Pedestes parnaca Fruhstorfer, 1910 : Deut. ent. Zeit. [Iris] 24:99. TL. Chiem Hoa, Tonkin. (♂, NHML)

[ THAILAND ]
Arnetta atkinsoni ; Godfrey,1930 : 368. (Prae: Me Song forest)
Arnetta atkinsoni ; Evans,1949 : 236. (Siam)
Arnetta atkinsoni ; Pinratana,1985 : 66, pl.22, figs.134,♂,♀. (Chiang Mai; Kanchanaburi; Chanthaburi)
Arnetta atkinsoni ; Yamaguchi & Aoki,2002 : 12, fig.17,♂. (Ubon Ratchathani: Phu Chong Na Yoi)
Arnetta atkinsoni ; Yamaguchi, Aoki & Akiyama,2005 : 44. (Ubon Ratchathani: Phu Chong Na Yoi)
Arnetta atkinsoni ; Katsuyama et al.,2006 : 44, fig.14,♂. (Nan: Ban Nam Poon)
Arnetta atkinsoni ; Ek-Amnuay,[2007] : 788, pl.362, figs.H160,♂,♂(Un),♂,♂(Un). (Kanchanaburi)
Arnetta atkinsoni ; Kimura et al.,2011 : 68, figs.♂,♂(Un),♂,♂(Un),♀,♀(Un). (Chiang Dao; Doi Suthep; Charng Phui; Thung Saleng Luang; Phu Khiewo; Erawan WF; Sai York Noi)
Arnetta atkinsoni ; Ek-Amnuay,2012 : 842, pl.389, figs.H156,♂,♂(Un),♂,♂(Un). (Kanchanaburi)
Arnetta atkinsoni ; S.Sophonviwatkul, C.Sunthornwiphat & T.Laola,2015- : Butterflies of Thailand, figs. (Chiang Mai; Phetchabun) (accessed: 2023.07.12)

[ LAOS ]
Arnetta atkinsoni ; Masui & Uehara,1996 : 5, pl.2, fig.25,♂,♂(Un). (Phong Sali)
Arnetta atkinsoni ; Osada, Uémura & Uehara,1999 : 224, pl.139, fig.♂(Un). (Boun Neua)
Arnetta atkinsoni ; Nakamura & Wakahara,2012 : 59.
Arnetta atkinsoni ; Onodera,2022 : 116, pl.253, figs.♂(dsf),♂(dsf,Un),♂(wsf),♂(wsf,Un),♀(wsf),♀(wsf,Un). (Oudomxai; Xaisomboun: Nam Phang)

[ VIETNAM ]
Pedestes parnaca ; Seitz,1927 : 1064. (Tonkin)
Arnetta atkinsoni ; Evans,1949 : 236. (Tonkin)
Pedestes parnaca ; Metaye,1957 : 78. (N)
Arnetta atkinsoni ; Inoué & Kawazoé,1966 ; 86-88, figs.73,♂,74,♂(Un). (Blao)
Arnetta atkinsoni ; Devyatkin & Monastyrskii,1999 : 162. (Ha Tinh; Do Son; Na Hang; Muong Nhe; Ba Be; Cuc Phuong; Ben En)
Arnetta atkinsoni ; Ikeda et al.,2001b : 65, figs.3:17,♂,18,♂(Un),5:28(♂genitalia). (Cuc Phuong)
Arnetta atkinsoni ; Luong et al.,2004 : 13, 87, figs.♀,♀(Un). (Cuc Phuong)
Arnetta atkinsoni ; Devyatkin & Monastyrskii,1999 : 163. (Ha Tinh; Do Son; Na Hang; Muong Nhe; Ba Be; Cuc Phuong; Ben En)
Arnetta atkinsoni ; Monastyrskii et al.,2006 : 45. (Thua Thien Hue: Nam Dong; A Luoi)
Arnetta atkinsoni ; Miyazaki, Saito & Saito,2007b : 10, figs.H-62,♂,♀. (Lam Dong: Dambri; Dai Duc Me; Madagui)
Arnetta atkinsoni ; Monastyrskii & Devyatkin,2015 : 75. (N / C / S)

[ THAILAND ]
Mae Hong Son : 4♂.

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Chiang Mai : 14♂ 1♀ (incl. photo).

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Phrae : 1♂.

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Tak : 1♀.

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[ LAOS ]
Oudomxai : 2♂ (photo by H.Onodera).

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The butterfly is locally common in secondary and montane forests at low to moderate elevations (Alt. 50 - 1000m) and is often found at forest edges and grasslands. Both sexes visit flowers and males occasionally come to the banks of streams and puddles to suck water.
Incredibly, I have seen natural hybridisation between this species and Iambrix salsala in Chiang Mai. This is the only case of mating I have ever seen between two different genera.

Moore, F.,1878 : Descriptions of new Asiatic Hesperidae.
Proc. zool. Soc. Lond. 1878(3):686-695, pl.45.

Moore, F.,1879 : A list of the lepidopterous insects collected by Mr.Ossian Limborg in upper Tenasserim, with descriptions of new species.
Proc. zool. Soc. Lond. 1878(4):821-858, pls. 51-53.

Fruhstorfer, H.,1910 : Neue Hesperiden des Indo-Malayischen Faunengebietes und Besprechung verwandter Formen.
Deut. ent. Zeit. [Iris] 24(3-5):58-104.

Huang, Z.F., Chiba, H., Jin, J., Kizhakke, A.G., Wang, M., Kunte, K. & Fan, X.L.,2019 :
A multilocus phylogenetic framework of the tribe Aeromachini (Lepidoptera: Hesperiidae: Hesperiinae), with implications for taxonomy and biogeography
Syst. Entomol. 44:163–178.

♂ (dsf), Chiang Mai, Thailand.	♀ (wsf), Chiang Mai, Thailand.
♂ (dsf), Chiang Mai, Thailand.	♂ (wsf), Phrae, Thailand.
Natural mating between Arnetta atkinsoni and Iambrix salsala, Chiang Mai, Thailand.

1999.09.04 - 2023.07.31.