♂ (wsf), Chanthaburi, Thailand.		♀ (wsf), Tak, Thailand.
♂ (dsf), Mae Hong Son, Thailand.		♀ (dsf), Chiang Mai, Thailand.
♂ (dsf), Phrae, Thailand.		♂ (dsf), Phetchaburi, Thailand.

ON : Pterygospidea ravi Moore, 1866 OD : Proc. zool. Soc. Lond. 1865(3):779. TL : Bengal. (NHML) Distribution : Nepal, India to Myanmar, Thailand, Laos, Cambodia, Vietnam, Yunnan. ● ssp.ravi ● ssp.atticus

Tagiades khasiana Moore, 1884 : J. asiat. Soc. Bengal, Part II. 53(1):51. TL. Khasi Hills, Assam. (♂, NHML)
Tagiades lugens Mabille, 1883 : C. r. Soc. ent. Belg. 27:lxxviii. TL. Saigon. (♂, NHML)
Tagiades khasiana epicharmus Fruhstorfer, 1910 : Deut. ent. Zeit. [Iris] 24:84. TL. Siam. (♀, NHML)

[ THAILAND ]
Tagiades atticus ; Godfrey,1916 : 142. (Chonburi: Dong Rek hills)
Tagiades khasiana epicharmus ; Godfrey,1916 : 143.
Tagiades ravi epicharmus ; Seitz,1927 : 1040. (Siam)
Tagiades atticus atticus f.atticus ; Godfrey,1930 : 360. (Prae: Me Song forest; Pak Jong; Muak Lek; Chonburi: Sriracha; Bangkok)
Tagiades atticus atticus f.khasiana ; Godfrey,1930 : 360. (Prae: Me Song forest; Pak Jong; Muak Lek; Chonburi: Sriracha; Bangkok)
Tagiades japetus ravi ; Evans,1949 : 135. (Siam)
Tagiades japetus ravi ; Shirôzu & Saigusa,1962 : 29, pl.1, fig.4,♀. (Doi Inthanon)
Tagiades japetus ravi ; Pinratana,1985 : 41, pl.15, figs.68a♀,♀(Un). (Nakhon Nayok; Nakhon Ratchasima)
Tagiades japetus ravi ; Yamaguchi & Aoki,2000 : 249, fig.10,♂. (Phu Pha Thaep; Ban Naudom; Phu Huay Sing; Ubon Ratchathani: Kaeng Tana)
Tagiades japetus ravi ; Yamaguchi & Aoki,2002 : 11. (Ubon Ratchathani: Phu Chong Na Yoi; Phu Hin Dang; Mukdahan: Phu Huay Sing; Ban Na Lak; Ban Gok Tum)
Tagiades japetus ravi ; Yamaguchi, Aoki & Akiyama,2005 : 43. (Ubon Ratchathani: Kaen Tana; Phu Chong Na Yoi; Phu Phung; Ban Ba Hai; Don Far Huan; Mukdahan: Ban Lao Kram; Ban Na Lak; Phu Hin Khan; Phu Ma Nom)
Tagiades japetus ravi ; Katsuyama et al.,2005 : 62, fig.8,♀. (Umphang: Tha Sai)
Tagiades japetus ravi ; Ek-Amnuay,[2007] : 762, pl.350, figs.H83a♀,♀(Un). (Trat)
Tagiades japetus ravi ; Kimura et al.,2011 : 45, figs.♂,♂(Un),♀,♀(Un),♂,♂(Un),♀,♀(Un). (Doi Suthep; Nakohn Nayok; Wan Nam Khiewo; Kao Soi Dao; Kratchin WF; Kao Khiewo; Mae Sohd; Erawan WF; Kuwai Yai; Ratchaburi)
Tagiades japetus ravi ; Ek-Amnuay,2012 : 814, pl.375, figs.H81a♀,♀(Un). (Trat)
Tagiades japetus ravi ; Okamoto et al.,2013 : 6. (Kanchanaburi: Mae Klong Station)
Tagiades japetus ravi ; S.Sophonviwatkul, C.Sunthornwiphat & T.Laola,2015- : Butterflies of Thailand, figs, Early Stages. (Sa Kaeo; Nakhon Nayok; Saraburi; Trat; Chanthaburi; Ratchaburi) (accessed: 2023.06.08)
Tagiades japetus ravi ; Chaianunporn & Chaianunporn,2019 : 85. (Chaiyaphum: Chulabhorn Dam)

[ LAOS ]
Tagiades japetus ravi ; Motono & Negishi,1989 : 79, pl.111, figs.B1,♂,B2,♂(Un),B3,♀,B4,♀(Un). (no loc.)
Tagiades japetus ravi ; Osada, Uémura & Uehara,1999 : 222, pl.136, figs.♂,♀. (Thakhek; Thateng)
Tagiades japetus ravi ; Nimura & Wakahara,2005 : 49. (Namdhua; Thabeng)
Tagiades japetus ravi ; Nakamura & Wakahara,2012 : 57.
Tagiades japetus ravi ; Onodera,2022 : 108, pl.244, figs.♂,♂(Un),♀,♀(Un),♀,♀(Un). (Bolikhamxai: Thabok; Houay Phet; Savannakhet)

[ CAMBODIA ]
Tagiades japetus ; Onodera,2007 : 29, fig.105,♂. (Siem Reap: Angkor)
Tagiades japetus ravi ; Kosterin,2019 : 99. (Kampot; Kep)
Tagiades japetus ravi ; Kosterin,2020 : 332. (Preah Vihear; Stueng Treng)
Tagiades japetus ravi ; Chartier & Kosterin,2022 : 125. (Kep; Koh Kong; Kampot; Preah Sihanouk; Preah Vihear; Siem Reap; Stueng Treng)

[ VIETNAM ]
Tagiades atticus ; Moulton,1923 : 134. (Pulo Condore)
Tagiades japetus sangareva ; Metaye,1957 : 77. (N)
Tagiades ravi epicharmus ; Metaye,1957 : 77. (S)
Tagiades japetus ravi ; Inoué & Kawazoé,1965b : 86-87, figs.33,♂,34,♂(Un),35,♀,36,♀(Un). (Trang Bom; Lai Thieu)
Tagiades japetus ravi ; Devyatkin & Monastyrskii,1999 : 161. (Bach Ma)
Tagiades japetus ; Vu,2003 : 58. (Phu Quoc)
Tagiades japetus ; Monastyrskii et al.,2006 : 44. (Thua Thien Hue: Bach Ma)
Tagiades japetus ravi ; Miyazaki, Saito & Saito,2007b : 7, figs.H-37,♂,♀. (Lam Dong: Dambri; Dai Duc Me; Dong Nai: Thac Mai; Bin Thuan: Dong Tien)
Tagiades japetus ravi ; Nihira,2009 : 101. (Phu Quoc)
Tagiades japetus ravi ; Monastyrskii & Devyatkin,2015 : 74. (C / S)
Tagiades japetus ravi ; Saito et al.,2017 : 76. (S: Nui Cam)

[ THAILAND ]
Mae Hong Son : 2♀.

Jan

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Dec

Chiang Mai : 8♂ 4♀ (incl. photo).

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Lampang : 1♂ (photo).

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Phrae : 1♂.

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Nan : 1♂.

Jan

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Tak : 1♂.

Jan

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Chonburi : 1♀.

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Chanthaburi : 2♀.

Jan

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Trat (Koh Chang dist.) : 2♂ 1♀.

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Phetchaburi : 1♂.

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[ LAOS ]
Vientiane : 1♂.

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The butterfly is very common in secondary and montane forests at low to moderate elevations (Alt. 30 - 1200m). Both sexes visit flowers and males occasionally come to the banks of streams and puddles to suck water. They often fly out from bushes along the trail and rest on the underside of leaves with their wings spread.

Zhang et al (2022:25) divided japetus into seven species. However, the number of taxa examined, especially those from continental regions, is inadequate, and there are also few differences in markings and male genitalia. They treated the Indian species as Tagiades ravi, Tagiades atticus atticus as the population of Thailand, and the population from W.Malaysia as Tagiades atticus balana. However, the type locality of atticus is considered to be W.Malaysia or peninsular Thailand. Furthermore, the type locality of balana is Borneo and is generally considered to be a Bornean subspecies. Thus, their treatment is rather questionable. I have therefore tentatively followed the traditional classification. Further taxonomic study is needed.

Moore, F.,1866 : On the lepidopterous insects of Bengal.
Proc. zool. Soc. Lond. 1865(3):755-822, 3 pls.

Mable, P.,1883 : Description d'hesperiens.
C. r. Soc. ent. belg. 27:li-lxxviii.

Moore, F.,1884 :
Descriptions of some New Asiatic Lepidoptera; chiefly from Specimens contained inthe Indian Museum, Calcutta.
J. asiat. Soc. Bengal, Part II. 53(1):16-52.

Fruhstorfer, H.,1910 : Neue Hesperiden des Indo-Malayischen Faunengebietes und Besprechung verwandter Formen.
Deut. ent. Zeit. [Iris] 24(3-5):58-104.

Zhang, J., Cong, Q., Shen, J. & Grishin, N.V.,2022 : Taxonoic changes suggested by the genomic analysis of Hesperiidae (Lepidoptera)
Insecta Mundi 0921: 1-135.

♂ (dsf), Lampang, Thailand.	♀ (dsf), Chiang Mai, Thailand.
♂, Chiang Mai, Thailand.	♂, Chiang Mai, Thailand.

1999.07.31 - 2024.04.30.