Cupha erymanthis erymanthis

Cupha erymanthis erymanthis (Drury, 1773)


♂, Chiang Mai, Thailand.		♀, Chiang Mai, Thailand.

♂, Satun, Thailand.		♂, Satun, Thailand.

♂, Yala, Thailand.		♂, Chiang Mai, Thailand.

♂, Vientiane, Laos.		♂, Vinh Phuc, Vietnam.

♂, Chonburi, Thailand.		♂ (ab), Phetchaburi, Thailand.

ON : Papilio erymanthis Drury, 1773
OD : Ill. nat. Hist. Exot. Insects 1: index, 29, pl.15, figs.3, 4(Un).
TL : "China".

Distribution : India to Myanmar, Thailand, Laos, Cambodia, Vietnam, Hainan, S.China, Taiwan, Langkawi, W.Malaysia, Singapore, Borneo, Sumatra, Palawan.

	● ssp.erymanthis
	● ssp.melanodotis

Synonyms
(--- The taxon marked with an asterisk is cited from other literature and I have not been able to confirm the original description.)

Papilio lotis Sulzer, 1776 : Gesch. Ins. nach linn. Syst :144, pl.16, fig.6. TL. Indien.
Cupha erymanthis palawana Fruhstorfer, 1900 : Stett. ent. Ztg. 60(4):344. TL. Palawan.
Cupha erymanthis nagara Fruhstorfer, 1912 : in Seitz, Grossschmett Erde 9:466. TL. 'West-Sumatra; Borneo'.
Cupha erymanthis form takamukui Matsumura, 1929 : Insecta Matsumurana 3(2/3):96, pl.4, fig.7,♂. TL. "Formosa: Hori" [Puli, Nantou, Taiwan]. (IFS)
Cupha erymanthis forma melan Esaki & Nomura, 1930 : Zephyrus 2(1):14, pl.3, figs.3♂, 4♂(Un). TL. "Taipei" [Taipei, Taiwan]. (IFS)
∗ "ab. decolorata Sonan, 1926 : Trans. nat. Hist. Soc. Formosa 16:181. TL. "Formosa" [Taiwan]." (Cited from Shirôzu,1960)
∗ "f. sadaichii Umeno, 1936 : Zephyrus 6:223." (Cited from Shirôzu,1960)

Records taken from Literature

[ THAILAND ]
Cupha erymanthis lotis ; Fruhstorfer,1906 : 319. (Siam)
Cupha erymanthis lotis ; Godfrey,1916 : 125.
Cupha erymanthis lotis ; Pendlebury,1923 : 35. (Nakhon Sri Thammarat: Khao Luang)
Cupha erymanthis lotis ; Godfrey,1927 : 123. (N)
Cupha erymanthis lotis ; Godfrey,1930 : 266. (all over)
Cupha erymanthis lotis ; Godfrey,1931 : 194. (Ko Tao)
Cupha erymanthis lotis ; Godfrey,1932a : 268. (P)
Cupha erymanthis lotis ; Pinratana,1979 : 8, figs.N10,♂,♂(Un). (Chiang Mai; Chonburi; Kanchanaburi; Nakhon Nayok; Nonthaburi; Petchaburi; Tak; Trang)
Cupha erymanthis lotis ; Pinratana & Eliot,1996 : 22, pl.5, fig.11,♂ (Chiang Mai; Tak; Nakhon Nayok; Kanchanaburi; Chonburi; Petchaburi; Ranong; Trang)
Cupha erymanthis lotis ; Yamaguchi & Aoki,2000 : 254, fig.82,♂. (Mukdahan: Ban Naudom)
Cupha erymanthis lotis ; Yamaguchi & Aoki,2002 : 24. (Mukdahan: Phu Huay Sing; Nam Tok Tad Ton; Ban Na Lak; Ban Gok Tum)
Cupha erymanthis lotis ; Yamaguchi, Aoki & Akiyama,2005 : 49. (Mukdahan: Ban Lao Kram; Nam Tok Tad Ton; Ubon Ratchathani: Phu Pung)
Cupha erymanthis lotis ; Katsuyama et al.,2005 : 69, fig.134,♂. (Umphang: Tha-Sai; Umphang)
Cupha erymanthis lotis ; Katsuyama et al.,2006 : 49, fig.101,♂. (Nan: Ban Nam Poon)
Cupha erymanthis lotis ; RDPB & DNP,2006-2008 : Butterflies in Hala-Bala Forest, fig.♂. (Narathiwat: Hala Bala) (last update: 2006.12.25 ; accessed: 2024.05.11)
Cupha erymanthis erymanthis ; Ek-Amnuay,[2007] : 356, pl.152, figs.N11,♂,♂(Un),♀. (Koh Samui; Kanchanaburi)
Cupha erymanthis erymanthis ; Les Day,2011-2022 : Samui Butterflies, figs.♂,♂(Un). (Koh Samui; Koh Phangan) (accessed: 2023.05.02)
Cupha erymanthis erymanthis ; Ek-Amnuay,2012 : 384, pl.165, figs.N214,♂,♂(Un),♀. (Koh Samui; Kanchanaburi)
Cupha erymanthis erymanthis ; Okamoto et al.,2013 : 9. (Kanchanaburi: Mae Klong Station)
Cupha erymanthis erymanthis ; S.Sophonviwatkul, C.Sunthornwiphat & T.Laola,2015- : Butterflies of Thailand, figs. (Khon Kaen; Sa Kaeo; Nakhon Ratchasima; Phetchaburi) (accessed: 2023.05.02)
Cupha erymanthis lotis ; Kimura et al.,2016 : 48, figs.♂,♂(Un),♀,♀(Un). (Sarika; Nakohn Nayork; Wan Nam Khiewo; Kratching WF; Kao Khiewo; Mae Sohd; Kwai Yai; Kaeng Krachan; Khlong Naka; Ka Chong; Than To WF.)
Cupha erymanthis erymanthis ; Chaianunporn & Chaianunporn,2019 : 86. (Chaiyaphum: Chulabhorn Dam)
Cupha erymanthis erymanthis ; Küppers & Janikorn,2020 : 30, figs.Abb.L-185-188. (Phang Nga)

[ LAOS ]
Cupha erymanthis lotis ; Motono & Negishi,1989 : 54, pl.41, figs.7,♂,8,♂(Un).(Vang Vieng; Pakse; Vientiane; Luang Prabang; Tha Ngone; Nam Ngum Dam)
Cupha erymanthis lotis ; Osada, Uémura & Uehara,1999 : 205, pl.47, fig.♂. (Xaignabouri)
Cupha erymanthis lotis ; Nimura & Wakahara,2005 : 41. (Namdhua)
Cupha erymanthis erymanthis ; Nakamura & Wakahara,2012 : 44.
Cupha erymanthis erymanthis ; Onodera,2022 : 26, pl.91, figs.♂,♂(Un). (Khammouan: Ban Lao)

[ CAMBODIA ]
Cupha erymanthis lotis ; Woodfield & Murton,2006 : figs,♂,♂(Un). (Botum Sakor NP.)
Cupha erymanthis ; Onodera,2007 : 28, fig.47♂. (Siem Reap: Phnum Kuhlen)
Cupha erymanthis ; Onodera,2008 : 58. (Mondolkiri: Bousra)
Cupha erymanthis erymanthis ; Kosterin,2019 : 97. (Koh Kong)
Cupha erymanthis erymanthis ; Kosterin,2020 : 326. (Siem Reap; Stueng Treng)
Cupha erymanthis erymanthis ; Chartier & Kosterin,2022 : 117. (Kep; Koh Kong; Mondulkiri; Preah Sihanouk; Ratanakiri; Siem Reap; Stueng Treng)

[ VIETNAM ]
Cupha erymanthis lotis ; Fruhstorfer,1906 : 319. (Tonkin / Annam)
Cupha lotis ; Vitalis,1919 : 226. (Annam)
Cupha dohertyi ; Lemée,1950 : 10. (Sontay)
Cupha erymanthis lotis ; Godfrey,1919 : 467. (S: Daban)
Cupha erymanthis lotis ; Metaye,1957 : 93. (N / C / S)
Cupha erymanthis erymanthis ; Ikeda et al.,2000 : 24, figs.1:7,♂,8,♂(Un), fig.9:3(♂,genitalia). (Cuc Phuong)
Cupha erymanthis ; Vu,2003 : 57. (Phu Quoc)
Cupha erymanthis ; Luong et al.,2004 : 5, 57, figs.♂,♂(Un). (Cuc Phuong)
Cupha erymanthis ; Monastyrskii et al.,2006 : 40. (Thua Thien Hue: Nam Dong; A Luoi; Huong Thuy; Phong Dien; Bach Ma)
Cupha erymanthis erymanthis ; Miyazaki, Saito & Saito,2006b : 17, fig.N-17,♂. (Lam Dong: Dambri; Dong Nai: Madagui; Thac Mai; Bin Thuan: Dong Tien)
Cupha erymanthis erymanthis ; Nihira,2009 : 99. (Phu Quoc)
Cupha erymanthis erymanthis ; Miyazaki et al.,2010 : 21, fig.N-17,♀. (Lam Dong: Dambri; Dai Duc Me)
Cupha erymanthis erymanthis ; Monastyrskii & Devyatkin,2015 : 32. (N / C / S)
Cupha erymanthis lotis ; Yokochi et al.,2015 : 40. (Khanh Hoa: Hon Ba)
Cupha erymanthis erymanthis ; Saito et al.,2017 : 81. (S: Nui Cam; Nui Ket)
Cupha erymanthis erymanthis ; Monastyrskii,2019 : 4(1):192-193, pl.25, fig.4a♂. (N: Na Hang; Tam Dao; Xuan Lien; Cat Ba; Cuc Phuong / C: Ben En; Pu Mat; Pu Hoat; Huong Son; Vu Quang; Phong Nha-Ke Bang)
Cupha erymanthis lotis ; Monastyrskii,2019 : 4(1):192-193. (C: Phong Dien; Bach Ma; Son Thanh; Ba To; Ngoc Linh; Kon Cha Rang; Kon Ka Kinh; Hon Ba; Nui Chua; Cat Loc; Yok Don; Binh Chau; Bu Gia Map; Chu Yang Sin / S: Cat Tien; Lo Go Sa Mat; Phu Quoc)

Specimens Examined

[ THAILAND ]
Mae Hong Son : 3♂ (incl. photo).

Jan Feb Mar Apr May Jun Jul Aug Sep Oct Nov Dec

Chiang Rai : 1♂ (photo).

Jan Feb Mar Apr May Jun Jul Aug Sep Oct Nov Dec

Chiang Mai : 12♂ 1♀ (incl. photo).

Jan Feb Mar Apr May Jun Jul Aug Sep Oct Nov Dec

Lamphun : 1♂ (photo).

Jan Feb Mar Apr May Jun Jul Aug Sep Oct Nov Dec

Lampang : 2♂ (photo).

Jan Feb Mar Apr May Jun Jul Aug Sep Oct Nov Dec

Phrae : 2♂ (photo).

Jan Feb Mar Apr May Jun Jul Aug Sep Oct Nov Dec

Nan : 4♂ (incl. photo).

Jan Feb Mar Apr May Jun Jul Aug Sep Oct Nov Dec

Chonburi : 1♂.

Jan Feb Mar Apr May Jun Jul Aug Sep Oct Nov Dec

Chanthaburi : 1♂.

Jan Feb Mar Apr May Jun Jul Aug Sep Oct Nov Dec

Trat : 4♂ (incl. photo by E.Ikari).

Jan Feb Mar Apr May Jun Jul Aug Sep Oct Nov Dec

Phetchaburi : 1♂.

Jan Feb Mar Apr May Jun Jul Aug Sep Oct Nov Dec

Ranong : 5♂.

Jan Feb Mar Apr May Jun Jul Aug Sep Oct Nov Dec

Surat Thani (Koh Phangan dist.) : 1♂.

Jan Feb Mar Apr May Jun Jul Aug Sep Oct Nov Dec

Satun : 7♂.

Jan Feb Mar Apr May Jun Jul Aug Sep Oct Nov Dec

Yala : 2♂ (incl. photo).

Jan Feb Mar Apr May Jun Jul Aug Sep Oct Nov Dec

[ LAOS ]
Vientiane : 1♂.

Jan Feb Mar Apr May Jun Jul Aug Sep Oct Nov Dec

[ VIETNAM ]
Vinh Phuc : 1♂.

Jan Feb Mar Apr May Jun Jul Aug Sep Oct Nov Dec

Lam Dong : 1♂.

Jan Feb Mar Apr May Jun Jul Aug Sep Oct Nov Dec

Kien Giang (Phu Quoc) : 3♂ (photo by E.Ikari).

Jan Feb Mar Apr May Jun Jul Aug Sep Oct Nov Dec

Notes

The butterfly is common in montane and secondary forests at low to moderate elevations (Alt. 10 - 1300m). Both sexes visit flowers and males occasionally come to the banks of streams and puddles to suck water.
From the Indo-China area to the Malay Peninsula, the species is highly variable and individual and seasonal variation seems to be more pronounced than geographical variation. There is a tendency for the ground color and markings become darker from north to south, but it is hard to separate them clearly. Thus I follow Tsukada (1985) and treat this as the nominotypical subspecies.

Literature

Drury, D.,[1773] : Illustrations of natural history; wherein are exhibited ...Illust. Nat. Hist. Exot. Insects
1:1-130, pl.1-50.

Sulzer, J.H.,1776 : Dr. Sulzers Abgekurzte Geschichte der Insecten nach dem Linaeischen System Gesch. Ins. nach linn. Syst.
(1):1-274, (2):pl.1-32.

Fruhstorfer, H.,1900 : Neue Cupha-Unterarten und Aufzählung der bekannten Species.
Stett. ent. Ztg. 60(4):343-347. (Apr.1900)

Fruhstorfer, H.,1912 : 6. Familie: Nymphalidae in Seitz, Gross-Schmett. Erde 9
457-488. (18.Mar.1912)

Matsumura, S.,1929 : New butterflies from Japan, Korea and Formosa
Insecta Matsumurana 3(2/3):87-107, pl.4.

Esaki, T. & Nomura, K.,1930 : Some new aberrant forms of the Formosan butterflies.
Zephyrus 2(1):11-14, pl.3.

Shirôzu, T.,1960 : Butterflies of Formosa in color.
Hoikusha, Osaka, Japan. [8] + 482 + [2]pp., 76 col.pls. (in Japanese)

Tsukada, E.,1985 : Butterflies of the South East Asian Islands. IV Nymphalidae (1).
Plapac Co., Ltd. 558 pp. 158.col.pls. (in Japanese)


♂, Yala, Thailand.	♂, Yala, Thailand.

♂, Chiang Mai, Thailand.	♂, Chiang Mai, Thailand.

2007.10.20 - 2024.05.31.