Charaxes bernardus hierax  Felder & Felder,[1867]

Charaxes bernardus hierax ♂ Up. Charaxes bernardus hierax ♂ Un. Charaxes bernardus hierax ♀ Up. Charaxes bernardus hierax ♀ Un.
♂, Tak, Thailand. ♀, Chiang Mai, Thailand.
Charaxes bernardus hierax ♂ Up. Charaxes bernardus hierax ♂ Un. Charaxes bernardus hierax ♀ Up. Charaxes bernardus hierax ♀ Un.
♂, Nan, Thailand. ♀, Nan, Thailand.
Charaxes bernardus hierax ♂ Up. Charaxes bernardus hierax ♂ Un. Charaxes bernardus hierax ♀ Up. Charaxes bernardus hierax ♀ Un.
♂, Ninh Binh, Vietnam. ♀, Vinh Phuc, Vietnam.
Charaxes bernardus hierax ♂ Up. Charaxes bernardus hierax ♂ Un. Charaxes bernardus hierax ♀ Up. Charaxes bernardus hierax ♀ Un.
♂, Phrae, Thailand. ♂, Phrae, Thailand.
Charaxes bernardus hierax ♂ Up. Charaxes bernardus hierax ♂ Un. Charaxes bernardus hierax ♀ Up. Charaxes bernardus hierax ♀ Un.
♂, Chiang Mai, Thailand. ♂, Chanthaburi, Thailand.
Charaxes bernardus hierax ♂ Up. Charaxes bernardus hierax ♂ Un. Charaxes bernardus hierax ♀ Up. Charaxes bernardus hierax ♀ Un.
♂, Nan, Thailand. ♀, Phrae, Thailand.
Charaxes bernardus hierax ♂ Up. Charaxes bernardus hierax ♂ Un. Charaxes bernardus hierax ♂ Up. Charaxes bernardus hierax ♂ Un.
♂, Vientiane, Laos. ♂, Kanchanaburi, Thailand.
Charaxes bernardus hierax ♂ Up. Charaxes bernardus hierax ♂ Un. Charaxes bernardus hierax ♂ Up. Charaxes bernardus hierax ♂ Un.
♂, Tak, Thailand. ♂, Tak, Thailand.
Charaxes bernardus hierax ♂ Up. Charaxes bernardus hierax ♂ Un. Charaxes bernardus hierax ♂ Up. Charaxes bernardus hierax ♂ Un.
♂, Nan, Thailand. ♂, Mae Hong Son, Thailand.
Charaxes bernardus hierax ♂ genitalia Charaxes bernardus hierax ♂ genitalia Charaxes bernardus hierax ♂ genitalia Charaxes bernardus hierax ♂ genitalia
♂ genitalia, Chaiyaphum, Thailand.

ON : Charaxes hierax Felder & Felder,[1867]
OD : Reise Fregatte Novara, Lep. Rhop. (3):442-443.
TL : Assam.

Distribution : Assam to Myanmar, Thailand, Laos, Vietnam, S.Yunnan.

Synonyms

Charaxes hipponax Felder & Felder,[1867] : Reise Fregatte Novara, Lep. Rhop. (3):443. TL. Assam.
Charaxes pleistoanax Felder & Felder,[1867] : Reise Fregatte Novara, Lep. Rhop. (3):443-444. TL. Assam.
Charaxes corax Felder & Felder,[1867] : Reise Fregatte Novara, Lep. Rhop. (3):444. TL. India septentrionalis.
Charaxes harpax Felder & Felder,[1867] : Reise Fregatte Novara, Lep. Rhop. (3):444-445. TL. no loc. [N.India]
Charaxes khasianus Butler,1872 : Lep. exot. : 98, pl.37, fig.6,(♂,♀) TL. Khasia Hills ; N.E.Bengal.
Charaxes agna Moore,[1879] : Proc. zool. Soc. Lond. 1878(4):832. TL. Moolai, Tenasserim.
Charaxes watti Butler,1880 : Proc. zool. Soc. Lond. 1880(2):148. TL. Upper Assam.
Charaxes hierax var.naganum Tytler,1911 : J. Bombay nat. Hist. Soc. 21(1):55. TL. Naga Hills.
Charaxes bernardus mahawedi Fruhstorfer,1914 : in Seitz, Grossschmett Erde. 9:731. TL. Than Moi ; Chiem Hoa, Tonkin ; Luang Prabang, Laos.
Charaxes polyxena hierax f.rossa Fruhstorfer,1914 : in Seitz, Grossschmett Erde. 9:732. TL. Assam (Khasia - Naga Hills).


Records taken from Literature

[ THAILAND ]
Charaxes polyxena agna ; Fruhstorfer,1914 : 732. (Siam)
Charaxes polyxena corax ; Godfrey,1916 : 133.
Charaxes polyxena hierax f.hierax ; Godfrey,1930 : 302-303. (Prae: Me Song forest / Chiang Rai / Me Wong)
Charaxes polyxena hierax f.agna ; Godfrey,1930 : 303. (Pak Jong / Muak Lek / Chonburi: Sriracha)
Charaxes polyxena agna ; Talbot,1936 ; 57.
Charaxes polyxena agna ; Davidson & Macbeth,1938 : 77.
Charaxes bernardus hierax ; Pinratana,1979 : 105, figs.N177a,♂,♂(Un),♂,♂,♀. (Chiang Mai / Chanthaburi / Kao Yai / Phitsanulok / Ratchaburi)
Charaxes marmax marmax ; Pinratana,1979 : figs.N178,♀. (in part)
Charaxes bernardus agna ; Pinratana,1979 : 105, not fig. (List up only.)
Charaxes aristogiton aristogiton ; Pinratana,1979 : 107, figs.N179,♂,♂(Un). (Chiang Mai / Chaiyaphum)
Charaxes kahruba ; Pinratana,1979 : fig.N180,♀. (in part)
Charaxes bernardus hierax ; Pinratana,1996 : 96, pl.70, figs.181a,♂(f.hierax),♂(f.hipponax),♂(f.pleistonax), pl.71,fig.181a,♀. (Chiang Mai / Chaiyaphum / Chonburi / Phisanulok / Chanthaburi / Nakhon Nayok / Kanchanaburi / Ratchaburi)
Charaxes marmax marmax ; Pinratana,1996 : pl.72, fig.182,♂. (in part)
Charaxes aristogiton aristogiton ; Pinratana,1996 : pl.72, figs.183,♂,♀. (in part)
Charaxes kahruba ; Pinratana,1996 : pl.73, fig.184,♀. (in part)
Charaxes bernardus hierax ; Yamaguchi & Aoki,2002 : 31, fig.84♂. (Mukdahan: Ban Na Lak)
Charaxes kahruba ; Yamaguchi & Aoki,2002 : 31, fig.85♀. (Mukdahan: Ban Na Lak)
Charaxes bernardus hierax ; Yamaguchi, Aoki & Akiyama,2005 : 52. 59, fig.22♂. (Ubon Ratchathani: Phu Chong Na Yoi / Mukdahan: Nam Tok Tad Ton)
Charaxes bernardus hierax ; Katsuyama et al.,2005 : 71, fig.176♂. (Umphang: Tha-Sai ; Umphang)
Charaxes bernardus hierax ; Katsuyama et al.,2006 : 50, fig.130♂. (Nan: Ban Nam Poon)
Charaxes bernardus hierax ; Ek-Amnuay,[2007] : 476,478, pl.212, figs.N194a,f.hierax♂,♂(Un),♂(f.paleistonax),♂(f.hipponax), ♀,♀(Un). (Wiang Pa Pao / Phu Khieo)
Charaxes bernardus crepax ; Ek-Amnuay,[2007] : pl.213, fig.N194b,♀(middle),♀(middle,Un). (in part)
Charaxes marmax marmax ; Ek-Amnuay,[2007] : pl.214, figs.N195,♂,♂(Un). (in part)
Charaxes aristogiton peridoneus ; Ek-Amnuay,[2007] : pl.214, figs.N196,♂,♂(Un),♂,♂(Un). (in part)
Charaxes kahrubax ; Ek-Amnuay,[2007] : pl.215, figs.N197,♀(middle),♀(middle,Un),♀(bottom,left). (in part)
Charaxes bernardus hierax ; Ek-Amnuay,2012 : 366, pl.156, figs.N197a,♂(f.hierax),♂(Un),♂(f.pleistoanax),♂(f.hipponax) ; pl.157,figs.197a,♀,♀(Un). (Wiang Pa Pao / Phu Khieo)
Charaxes bernardus crepax ; Ek-Amnuay,2012 : pl.157, fig.N197b,♀(middle),♀(middle,Un). (in part)
Charaxes aristogiton aristogiton ; Ek-Amnuay,2012 : pl.158, figs.N199,♂(top,left),♂(top,center),♂(top,Un). (in part)
Charaxes kahruba ; Ek-Amnuay,2012 : pl.159, figs.N200,♀(middle,left),♀(middle,Un). (in part)
Charaxes bernardus hierax ; Okamoto et.al.,2013 : 9. (Kanchanaburi: Mae Klong Station)
Charaxes bernardus hierax ; S.Sophonviwatkul, C.Sunthornwiphat & T.Laola,2015- : Butterflies of Thailand, figs, Early Stages. (Sa Kaeo / Saraburi / Chanthaburi / Nakhon Nayok / Phetchabun)
Charaxes bernardus hierax ; Kimura et al.,2016 : 130, figs.♂,♂(Un),♂,♂(Un),♂,♂(Un),♂,♂(Un),♂,♂(Un),♂,♂(Un),♀,♀(Un),♀,♀(Un). (Chiang Dao / Doi Suthep / Doi Inthanon / Khun Lao / Doi Pa Chang / Thung Saleng Luang / Phu Khiewo / Sarika / Kao Yai / Nakohn Nayok / Wan Nam Khiewo / Bung Khan / Kao Soi Dao / Kao Khiewo / Sai York Noi / Kaeng Krachan)

[ LAOS ]
Charaxes polyxena mahawedi ; Fruhstorfer,1914 : 731. (Luang Prabang)
Charaxes polyxena f.mahawedi ; Dubois & Vitalis de Salvaza,1924 : 47. (Laos; Luang-Prabang)
Charaxes polyxena f.corax ; Dubois & Vitalis de Salvaza,1924 : 47. (Laos)
Charaxes polyxena f.hipponax ; Dubois & Vitalis de Salvaza,1924 : 47. (Luang-Prabang)
Charaxes polyxena f.khasianus ; Dubois & Vitalis de Salvaza,1924 : 47. (Thado)
Charaxes bernardus hierax ; Motono & Negishi,1989 : 63, pl.79, figs.1,2. (Vang Vieng / Phatong)
Charaxes aristogiton aristogiton ; Motono & Negishi,1989 : 63, pl.79, figs.5,6. (Vang Vieng)
Charaxes bernardus hierax ; Osada, Uémura & Uehara,1999 : 208, pl.82, figs.♂,♀,♂,♂,♂,♂(Un),♂(Un),♂(Un),♀. (Lak Sao / Xaignabouri / Thabok / Nam Tha)
Charaxes bernardus hierax ; Nimura & Wakahara,2005 : 43. (Thabok / Namdhua)
Charaxes bernardus hierax ; Onodera,2009b : 35,39,41,42,43. (Vientiane: Namsong Riverside / Bolikhamxai: Thabok; Phon Si / Khammouan: Nacay)
Charaxes aristogiton aristogiton ; Osada, Uémura & Uehara,1999 : 209, pl.83, figs.♂,♂(Un),♀. (Thabok / Vang)
Charaxes bernardus hierax ; Nakamura & Wakahara,2012 : 47.
Charaxes aristogiton aristogiton ; Nakamura & Wakahara,2012 : 47.

[ CAMBODIA ]
Charaxes bernardus hierax ; Woodfield & Murton,2006 : fig.♀. (Botum Sakor NP.)
Charaxes kahruba ; Woodfield & Murton,2006 : fig.♀. (Botum Sakor NP.)

[ VIETNAM ]
Charaxes hipponax ; Joannis,1901 : 319. (Tonkin: Cao Bang)
Charaxes polyxena mahawedi ; Fruhstorfer,1914 : 731. (Tonkin: Than Moi ; Chiem Hoa)
Charaxes mahawedi ; Dubois & Vitalis de Salvaza,1919 : 233. (Tonkin)
Charaxes polyxena f.mahawedi ; Dubois & Vitalis de Salvaza,1924 : 47. (Tonkin: Chiem-Hoa)
Charaxes polyxena f.khasianus ; Dubois & Vitalis de Salvaza,1924 : 47. (Lao-Kay)
Charaxes polyxena mahawedi ; Metaye,1957 : 99. (N)
Charaxes polyxena mahawedi f.khasianus ; Metaye,1957 : 99. (N)
Charaxes marmax ; Ikeda et al.,2000 : figs.8:9♂,10♂(Un). (Cuc Phuong) (in part)
Charaxes bernardus mahawedi ; Ikeda et al.,2000 : 37, figs.8:12♂,13♂,14♂,15♂(Un), fig.9:48(♂,genitalia). (Cuc Phuong)
Charaxes bernardus hierax ; Hao et al.,2004 : 8, 72, figs.♂,♂(Un). (Cuc Phuong)
Charaxes aristogiton peridoneus ; Hao et al.,2004 : 8, 72, figs.♂,♂(Un). (Cuc Phuong)
Charaxes bernardus hierax ; Miyazaki, Saito & Saito,2006b : 35, figs.N-116♂,♂,♀. (Lam Dong: Dambri / Dong Nai: Thac Mai / Bin Thuan: Dong Tien)
Charaxes bernardus ; Monastyrskii et al.,2006 : 42. (Thua Thien Hue: Nam Dong; Huong Thuy; Phong Dien; Bach Ma)
Charaxes aristogiton peridoneus ; Miyazaki, Saito & Saito,2007c : 39, fig.N-123♀. (Lam Dong: Madagui)
Charaxes bernardus hierax ; Monastyrskii & Devyatkin,2015 : 42. (N / C / S)
Charaxes bernardus hierax ; Yokochi et al.,2015 : 40, figs.49,♀,49v,♀(Un). (Khanh Hoa: Hon Ba)


Specimens Examined

[ THAILAND ]
Mae Hong Son : 10♂ 2♀ (incl.photo).
JanFebMarAprMayJun JulAugSepOctNovDec

Chiang Mai : 22♂ 6♀ (incl.photo).
JanFebMarAprMayJun JulAugSepOctNovDec

Lampang : 2♂ 1♀ (incl.photo).
JanFebMarAprMayJun JulAugSepOctNovDec

Phrae : 7♂ 8♀.
JanFebMarAprMayJun JulAugSepOctNovDec

Nan : 12♂ 2♀.
JanFebMarAprMayJun JulAugSepOctNovDec

Loei : 1♂ (photo).
JanFebMarAprMayJun JulAugSepOctNovDec

Nongkai : 1♂.
JanFebMarAprMayJun JulAugSepOctNovDec

Tak : 13♂.
JanFebMarAprMayJun JulAugSepOctNovDec

Kanchanaburi : 3♂.
JanFebMarAprMayJun JulAugSepOctNovDec

Phetchabun : 5♂ (incl.photo).
JanFebMarAprMayJun JulAugSepOctNovDec

Chaiyaphum : 2♂.
JanFebMarAprMayJun JulAugSepOctNovDec

Sriracha : 1♀.
JanFebMarAprMayJun JulAugSepOctNovDec

Chanthaburi : 3♂ 2♀.
JanFebMarAprMayJun JulAugSepOctNovDec

Trat : 1♂.
JanFebMarAprMayJun JulAugSepOctNovDec

Phetchaburi : 3♂.
JanFebMarAprMayJun JulAugSepOctNovDec


[ LAOS ]
Luang Phabang : 1♀ (photo by H.Onodera).
JanFebMarAprMayJun JulAugSepOctNovDec

Bolikhamxai : 1♀ (photo by H.Onodera).
JanFebMarAprMayJun JulAugSepOctNovDec

Khammouan : 1♀ (photo by H.Onodera).
JanFebMarAprMayJun JulAugSepOctNovDec

Vientiane : 4♂.
JanFebMarAprMayJun JulAugSepOctNovDec


[ VIETNAM ]
Vinh Phuc : 1♀.
JanFebMarAprMayJun JulAugSepOctNovDec

Ninh Binh : 1♂.
JanFebMarAprMayJun JulAugSepOctNovDec


Notes
The butterfly is very common in montane forest. Both male and female are polymorphic and very variable. Some examples from W.Thailand (Tak prov.) are very similar to ssp.crepax.

Literature

Felder, C. & R. Felder,1865-1875 :
Rhopalocera. In Reise der Osterreichischen Fregatte "Novara" um die Erde in den Jahren 1857, 1858, 1859 unter den Befehlen Commodore B. von Wullerstorf-Urbair. Zoologischer Theil. Zweiter Band:
Abtheilung. Wien : vi + 548 + [1] pp in 5 parts, 140 pls.

Butler, A.G.,1869-1874 : Lepidoptera exotica, or Descriptions an Illustrations of Exotic Lepidoptera.
London, E.W.Janson 4to:[vi]+190+[vi] pp, pls,1-64 (20 parts)

Moore, F.,[1879] :
A list of the lepidopterous insects collected by Mr.Ossian Limborg in upper Tenasserim, with descriptions of new species.
Proc. zool. Soc. Lond. 1878(4):821-859, 3 pls.

Butler, A.G.,1880 : On new and little-known butterflies from India.
Proc. zool. Soc. Lond. 1880(2):147-152, pl,15.

Tytler, H.,1911 : Notes on butterflies from the Naga Hills.
J. Bombay nat. Hist. Soc. 21(1):48-65, pl.A.

Charaxes bernardus hierax ♂ Charaxes bernardus hierax ♀
♂, Chiang Mai, Thailand. ♀, Chiang Mai, Thailand.
Charaxes bernardus hierax ♂ Charaxes bernardus hierax ♀
♂, Chiang Mai, Thailand. ♀, Chiang Mai, Thailand.
Charaxes bernardus hierax ♂ Charaxes bernardus hierax ♂
♂, Chiang Mai, Thailand. ♂, Chiang Mai, Thailand.
Charaxes bernardus hierax ♂ Charaxes bernardus hierax ♂
♂, Chiang Mai, Thailand. ♂, Chiang Mai, Thailand.
Charaxes bernardus hierax ♂ Charaxes bernardus hierax ♂
♂, Phetchabun, Thailand. ♂, Phetchabun, Thailand.

2006.12.09 - 2019.09.30.

A Check list of Butterflies in Indo-China, Chiefly from THAILAND, LAOS & VIETNAM. Copyright © Y.Inayoshi.